Liu, Dong https://orcid.org/0009-0003-1502-5657
Chang, Ruiying
Tang, Haiping https://orcid.org/0000-0002-9599-8162
Funding for this research was provided by:
Second Tibetan Plateau Scientific Expedition and Research Program (STEP)
National Natural Science Foundation of China (41977398)
National Natural Science Foundation of China (42271123)
Youth Innovation Promotion Association of the Chinese Academy of Sciences (Y2022096)
Article History
Received: 10 March 2025
Accepted: 24 December 2025
First Online: 6 January 2026
Competing interests
: The authors declare no competing interests.