Wu, Jiapeng
Zhang, Baoshan
Ma, Yuexi
Kuang, Chunyi
Hong, Yiguo
Funding for this research was provided by:
National Natural Science Foundation of China (42476141)
National Natural Science Foundation of China (42276130)
Open Fund of Hainan Xisha Marine Environment National Observation and Research Station (2025001)
Graduate Innovative Research Grant Program of Guangzhou Education Bureau (2024312281)
Basic and Applied Basic Research Foundation of Guangdong Province (2023B1515120029)
Science and Technology Projects in Guangzhou (2025A03J3103)
Article History
Received: 14 October 2025
Accepted: 22 June 2026
First Online: 29 June 2026
Competing interests
: The authors declare no competing interests.