Jiang, Shuhong
Li, Shuo
Han, Shenglei
Li, Chen
Cao, Xuwen
Liu, Changlin
Miao, Chunlong
Dong, Caichao
Zhang, Yanming
Liu, Kaiqiang
Liu, Yuyan
Wang, Qian
Wang, Hong-yan
Shao, Changwei
Funding for this research was provided by:
the Taishan Scholars Program (tsqnz20250759)
the Taishan Scholars Program (tstp20221149)
the Central Public-interest Scientific Institution Basal Research Fund, YSFRI, CAFS (NO. 20603022025008)
National Key R&D Program of China (2024YFD2400901)
National Key R&D Program of China (2024YFD2400901)
the Key Research and Development Project of Shandong Province (2025CXGC010802)
the Key Research and Development Project of Shandong Province (2024LZGC005)
the AoShan Talents Cultivation Program Supported by Qingdao National Laboratory for Marine Science and Technology (2017ASTCP-ES06)
the National Ten-Thousands Talents Special Support Program
the Improvement of Aquaculture Seed Industry Program of Yantai (2024)
the Central Public-interest Scientific Institution Basal Research Fund, CAFS (2023TD19)
Article History
Received: 19 January 2026
Accepted: 26 June 2026
First Online: 4 August 2026
Ethics
Competing interests: The authors declare no competing interests.