Shang, Huiying
Huyan, Yang
Lu, Yuan
Guo, Ming
Li, Bin
Funding for this research was provided by:
the Basic Research Program of the Shaanxi Academy of Sciences (Nos. 2022K-13, and 2024p-05)
the Basic Research Program of the Shaanxi Academy of Sciences (No. 2023K-14)
the Key R&D Program of the Shaanxi Provincial Department of Science and Technology (No. 2025NC-YBXM-057)
the National Natural Science Foundation of China (No. 32201588)
he Shaanxi Changqing National Nature Reserve Administration's 2022 Central Financial National Park Subsidy Project
Article History
Received: 7 November 2025
Accepted: 7 July 2026
First Online: 14 July 2026
Competing interests
: The authors declare no competing interests.