Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (19H04937)
MEXT | Japan Society for the Promotion of Science (17H05977)
MEXT | Japan Society for the Promotion of Science (17H05575)
MEXT | Japan Society for the Promotion of Science (17K14939)
MEXT | Japan Society for the Promotion of Science (19H05225)
MEXT | Japan Society for the Promotion of Science (18H05137)
MEXT | Japan Society for the Promotion of Science (17K19462)
MEXT | Japan Society for the Promotion of Science (16H04656)
MEXT | Japan Society for the Promotion of Science (16H01279)
MEXT | JST | Precursory Research for Embryonic Science and Technology (JPMJPR1882)
Takeda Science Foundation
Uehara Memorial Foundation
Naito Foundation
The Nakajima Foundation
Toray Science Foundation
Article History
Received: 16 July 2019
Accepted: 13 January 2020
First Online: 29 January 2020
Competing interests
: The authors declare no competing interests.