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Article History

Received: 18 April 2024

Accepted: 30 October 2024

First Online: 7 November 2024

Declarations

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: The authors declare no competing interests. Yang Meng, Qiang Xu, Yanling Zhang, Aiguo Wang, Jianwei Wang and Weimin Guo were employed by Zhengzhou Tobacco Research Institute of CNTC (Zhengzhou, China). Guangqing Chen, Jianjun Liu and Shuoye Zhou were employed by Henan Provincial Tobacco Company (Zhengzhou, China). Ding Yan and Xianjie Cai were employed by Shanghai Tobacco Company (Shanghai, China). Junying Li and Xuchu Chen were employed by Pingdingshan Branch of Henan Provincial Tobacco Company (Pingdingshan, China). Qiuying Li and Qiang Zeng were employed by Nanping Branch of Fujian Provincial Tobacco Company (Nanping, China). The remaining authors declare that the research was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest.

: The authors declare that the collection of plant material (tobacco leaves) and the experimental studies complied with the relevant institutional, national and international guidelines and legislation. The authors confirm that all methods were carried out in accordance with relevant guidelines in the method section. The authors comply with the IUCN Policy Statement on Research Involving Species at Risk of Extinction and the Convention on the Trade in Endangered Species of Wild Fauna and Flora. The authors ensure that the collection of tobacco leaves samples had been licensed by the local tobacco production regulatory agency.