Sato, Takehiro
Kubo, Daisuke
Hirasawa, Yu
Yoneda, Minoru
Kimura, Ryosuke
Tajima, Atsushi
Kato, Hirofumi
Funding for this research was provided by:
Japan Society for the Promotion of Science (19H01356, 16H01954)
Article History
Received: 8 July 2025
Accepted: 22 September 2025
First Online: 27 October 2025
Declarations
:
: The authors declare no competing interests.
: Permissions for the excavation of the Hamanaka 2 site were granted by the Board of Education of Hokkaido. Permissions for subsequent scientific analyses, including genetic studies, were granted by the Centre for Ainu and Indigenous Studies, Hokkaido University, which is temporarily storing these remains for archaeological and anthropological research. We consulted with the Ainu Association of Hokkaido regarding these studies, and the whole-genome analysis was approved by the ethics committee of Kanazawa University.