Xiao, Fangjing
Cui, Xiaomei
Jiang, Guibin
Bu, Duo
Zhang, Qiangying
Funding for this research was provided by:
the National Natural Science Foundation of China (No. 22266032, 22466032)
Central Financial Support Special Funds for Local Colleges and Universities (No. [2024]01, [2025]01)
Science and Technology projects of Xizang Autonomous Region (XZ202501ZY0072, XZ202501ZY0008)
Talent Incentive Project of Tibet University (No. ZDRC202410)
Science and Technology Plan Projects of Shigatse City (No. RKZ2024ZY-06)
the Lhasa Municipal Major Science and Technology Project (No. LSKJ202543)
Article History
Received: 20 June 2025
Accepted: 29 September 2025
First Online: 4 November 2025
Competing interests
: The authors declare no competing interests.