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Authors
Sun, Qi
Wu, Lei
Zhao, Ji-Min
Zhang, Yan-Wen
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Funding
Funding for this research was provided by:
Changchun Normal University (CSJJ2023008ZK, CSJJ2023008ZK)
National Natural Science Foundation of China (32272757, 32272757)
Liaoning province science and technology plan-combined plan (2024JH2/102600132)
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Article History
Received: 5 June 2025
Accepted: 21 October 2025
First Online: 24 November 2025
Declarations
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: The authors declare no competing interests.
: All plant materials were used in accordance with national and international standards and local laws and regulations. The use of all plant materials does not pose any risk to other species in nature. No specific permission is required for the collection of all samples described in this study. The specimens of the research species were kept in the herbarium of Liaodong University, No. CP2002-3, and the appraiser by Professor Zhang Yanwen.
