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Authors
Twesigomwe, David
Aitman, Timothy J.
Wilson, James F.
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Funding
Funding for this research was provided by:
Sydney Brenner Charitable Trust
Chief Scientist Office of the Scottish Government Health Directorates (SGP/1)
Chief Scientist Office of the Scottish Government (CZB/4/276 and CZB/4/710)
The Medical Research Council Whole Genome Sequencing for Health and Wealth Initiative (MC/PC/15080)
The MRC Human Genetics Unit quinquennial programme “QTL in Health and Disease” (U. MC_UU_00007/10)
Arthritis Research UK and the European Union framework program 6 EUROSPAN project (LSHG-CT-2006-018947)
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Article History
Received: 25 March 2025
Accepted: 28 October 2025
First Online: 26 November 2025
Declarations
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: The authors declare no competing interests.
: The Orkney Complex Disease Study (ORCADES) was supported by the Chief Scientist Office of the Scottish Government (CZB/4/276, CZB/4/710), a Royal Society URF to J.F.W., the MRC Human Genetics Unit quinquennial programme “QTL in Health and Disease”, Arthritis Research UK and the European Union framework program 6 EUROSPAN project (contract no. LSHG-CT-2006-018947). The Viking Health Study – Shetland (VIKING I) was supported by the MRC Human Genetics Unit quinquennial programme grant “QTL in Health and Disease”. We acknowledge the MRC Human Genetics Unit programme grant, “Quantitative traits in health and disease” (U. MC_UU_00007/10). The Scottish Genomes Partnership was funded by the Chief Scientist Office of the Scottish Government Health Directorates (SGP/1) and The Medical Research Council Whole Genome Sequencing for Health and Wealth Initiative (MC/PC/15080). DT is supported by funding from the Sydney Brenner Charitable Trust.
