Liu, Xiaoxuan
An, Ran
Wang, Guoyun
Funding for this research was provided by:
National Key Research and Development Program of China (2022YFC2704000)
Special foundation for Taishan Scholars (No. tstp20230657)
Major Basic Research of Natural Science Foundation of Shandong (ZR2021ZD34)
National Natural Science Foundation of China (grant numbers 82371653 and 82071621)
General Project of Shandong Natural Science Foundation (ZR2023MH177)
Article History
Received: 21 February 2025
Accepted: 29 October 2025
First Online: 27 November 2025
Competing interests
: The authors declare no competing interests.
: The study protocol was approved by the NCHS Ethics Review Board, and all participants provided informed consent.