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Article History

Received: 2 February 2025

Accepted: 23 April 2025

First Online: 28 April 2025

Change Date: 11 July 2025

Change Type: Update

Change Details: The original online version of this Article was revised: The original version of this Article contained an error in the Acknowledgements section. It now reads: “The authors thank Dr. Atsuo Ogura at Bioresource Engineering, RIKEN, for help with in vitro fertilization of <i>Slc12a2</i>^<i>Em1</i> mice; Dr. Atsushi Toyoda at Comparative Genomics Laboratory, National Institute of Genetics, for technical support with RNA-seq; and Dr. Mikio Furuse of the National Institute for Physiological Sciences for providing rabbit anti-Claudin 9 antiserum. This study was supported by grants to HM from the Japan Society of Promotion of Sciences (KAKENHI; grant numbers 18K0933, 21K09598, and 24K12688), and from the JSPS Platform for Advanced Genome Science (grant number 212118), by a grant to TM from the Uehara Memorial Foundation (grant number 201820479), and by a grant to SI from the Amgen Foundation (Undergraduate Summer Research Program in Science and Biotechnology 2022).”

Declarations

:

: The authors declare no competing interests.

: All experimental protocols were approved by Institutional Animal Care and Use Committee of NHO Tokyo Medical Center (23-animal-01, 18-animal-02). All experiments were performed in accordance with relevant guidelines and regulations. Authors complied with the ARRIVE guidelines ().