Murakami, Yuki
Hosomi, Ryota
Tanaka, Genki
Murakami, Hirokazu
Kanto, Ayaka
Kimura, Takahiro
Imamura, Yukio
Yoshida, Munehiro
Fukunaga, Kenji
Funding for this research was provided by:
Japan Society for the Promotion of Science (20K21587)
Japan Society for the Promotion of Science (20K21587)
Toyo Suisan Foundation
Tojuro Iijima Foundation for Food Science and Technology
the Kansai University Organization for Research and Development of Innovative Science and Technology
Article History
Received: 11 June 2025
Accepted: 30 January 2026
First Online: 13 February 2026
Competing interests
: The authors declare no competing interests.
: The study was conducted in accordance with relevant guidelines and regulations, such as Act on Welfare and Management of Animals and Guidelines for Proper Conduct of Animal Experiments in Japan and the experimental protocols approved by the Animal Ethics Committee of the Kansai University, Suita, Japan (approval number: 2205; approval date: 24/3/2021). Additionally, all animal experiments were carried out in accordance with ARRIVE guideline ( ).