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Authors
Mizuno, Shuntaro
Ishida, Hiroho
Konno, Ryou
Teshima, Natsuko
Yokota, Tomoko
Khin, Shwe Thiri Maung Maung
Takemae, Hitoshi
Oi, Toru
Fukuda, Fujiko
Masuda, Tsuneyuki
Kikkawa, Yasuhiro
Oka, Tomoichiro
Aihara, Naoyuki
Shiga, Takanori
Kamiie, Junichi
Murakami, Hironobu
Mizutani, Tetsuya
Nagai, Makoto
Oba, Mami https://orcid.org/0000-0002-4187-3217
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Funding
Funding for this research was provided by:
JSPS KAKENHI (21K05947)
JSPS Program for Forming Japan’s Peak Research Universities（J-PEAKS） (JPJS00420230003)
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Article History
Received: 5 November 2025
Accepted: 23 March 2026
First Online: 9 April 2026
Declarations
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: The authors declare no competing interests.
: The present study was carried out according to the Fundamental Guidelines for Proper Conduct of Animal Experiments and Related Activities in Academic Research Institutions under the jurisdiction of the Ministry of Education, Culture, Sports, Science and Technology of Japan. This study is reported in accordance with ARRIVE guidelines ( ). All pathogen- and animal-related experiments were reviewed and approved by the Safety Management Regulations for Pathogens Committee of Azabu University (Approval No. 21–13) and the Animal Experiment Committee of Azabu University (Approval No. 240213–4), respectively, and were conducted in accordance with the Azabu University Safety Management for Pathogens Implementation Regulations and Manual, as well as the Azabu University Animal Experiment Implementation Regulations and Manual.
