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Article History

Received: 26 September 2025

Accepted: 23 March 2026

First Online: 1 April 2026

Declarations

:

: The authors declare no competing interests.

: This study was conducted following the 2012 International Guidelines for Biomedical Research Involving Animals (Council for International Organizations of Medical Sciences, ), the care and use of laboratory animals were in full compliance with these guidelines. Furthermore, the study species is not listed in the List of Protected Animals in China and experimental research on this species is legal in China, no institutional permission was required for the collection of animals in this study.The experimental animals in this study were slaughtered at the dedicated slaughterhouse of Jiangquan Agriculture and Animal Husbandry Co., Ltd. in Linyi City, Shandong Province. The slaughter was performed by a colleague holding a veterinary qualification certificate after the animals were euthanized by electrocution. Our study has taken the informed consent of the animal owner. All animal care and treatment procedures were approved by Ethics Committee of Shandong Agricultural University, China, and performed according to the Committee’s guidelines and regulations (Approval No.: 2004006). We declare that this study is reported by ARRIVE guidelines.All authors have read and approved this version of the article, and due care has been taken to ensure the integrity of the work. No part of this paper has been published or submitted elsewhere. No conflict of interest exits in the submission of this manuscript.