Rahmaninia, Hossein
Ejtehadi, Hamid
Memariani, Farshid
Abedi, Mehdi
Palacio, Sara
Funding for this research was provided by:
Office of the Vice-President for Research and Technology of Ferdowsi University of Mashhad (3/60075)
European Union’s Horizon 2020 (H2020-MSCA RISE-777803)
Article History
Received: 20 October 2025
Accepted: 10 April 2026
First Online: 18 April 2026
Declarations
:
: The authors declare no competing interests.
: All field studies and collection of wild plant material were performed in accordance with relevant institutional, national, and international guidelines under formal permission from the Iranian Natural Resources and Watershed Management Organization. Voucher specimens were deposited at the FUMH herbarium under accession numbers included in the manuscript.