Yuan, Hang
Li, Junjie
Guan, Bo
Chu, Guangdi
Jiao, Wei
Zheng, Hongzhi
Liu, Xingchi
Zhao, Jianchang
Li, Jinhua
Li, Jianmin
Yang, Xuecheng
Niu, Haitao
Funding for this research was provided by:
the National Natural Science Foundation of China (52475027)
the National Natural Science Foundation of China (52525504)
the Natural Science Foundation of Shandong Province (ZR2024MH105)
the Shandong Provincial Key Research and Development Program (2022CXGC020505)
the National Key Research and Development Program of China (2023YFC2013405)
Article History
Received: 18 May 2025
Accepted: 11 December 2025
First Online: 13 January 2026
Competing interests
: The authors declare no competing interests.