Xiao, Lushan
Liang, Shengxing
Zeng, Lin
Cai, Shumin
Wang, Jiaren
Hong, Chang
Li, Yan
Li, Ruining
Jiang, Pu
Xie, Zebin
Li, Ting
Wu, Shanshan
Liu, Li
Wu, Gongfa
Lai, Weinan
Funding for this research was provided by:
National Nature Science Foundation of China (82404077)
National Nature Science Foundation of China (82400664)
National Nature Science Foundation of China (82172751)
Guangdong Natural Science Foundation (2022A1515110656)
Guangdong Province College Students' Innovative Entrepreneurial Training Progarm (S202412121153)
the Postdoctoral Fellowship Program of CPSF (GZC20240663)
National Key R&D Program of China (2021YFF1201304)
Article History
Received: 28 October 2025
Accepted: 22 December 2025
First Online: 6 January 2026
Competing interests
: The authors declare no competing interests.