Imaizumi, Riki https://orcid.org/0000-0002-9578-2513
Waki, Toshiyuki https://orcid.org/0000-0002-0128-096X
Hattori, Yoshikazu https://orcid.org/0000-0002-0013-5584
Takeshita, Kohei https://orcid.org/0000-0003-3469-9844
Sumita, Miru https://orcid.org/0009-0002-6300-1038
Kawaguchi, Kazutomo
Kumeta, Hiroyuki https://orcid.org/0000-0003-3713-2122
Umeda, Kenichi https://orcid.org/0000-0002-3650-3828
Sato, Kyohei
Yanai, Taro https://orcid.org/0009-0001-2019-3271
Matsuura, Hiroaki
Yokoyama, Takeshi
Omura, Risa
Nakatani, Kayo
Sakai, Naoki https://orcid.org/0000-0001-8544-1187
Kawagiwa, Yukimura
Doi, Yamato
Yasuda, Aoi
Nakano, Takuya
Uno, Kaichi
Yoshida, Kunihiro
Tsunashima, Misato
Saio, Tomohide https://orcid.org/0000-0003-3639-7399
Tanaka, Yoshikazu https://orcid.org/0000-0002-7893-3236
Takahashi, Seiji https://orcid.org/0000-0002-2288-4340
Kodera, Noriyuki https://orcid.org/0000-0003-4880-8423
Kataoka, Kunishige https://orcid.org/0000-0001-7096-7565
Yamamoto, Masaki https://orcid.org/0000-0002-1311-1768
Yamashita, Satoshi https://orcid.org/0000-0003-2978-3216
Nakayama, Toru https://orcid.org/0000-0002-5275-4731
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (23H05470)
MEXT | Japan Society for the Promotion of Science (18H03938)
MEXT | Japan Society for the Promotion of Science (21J22186)
MEXT | Japan Society for the Promotion of Science (22KJ1466)
MEXT | Japan Society for the Promotion of Science (24K23286)
MEXT | Japan Society for the Promotion of Science (20K15448)
MEXT | Japan Society for the Promotion of Science (25H01418)
MEXT | Japan Society for the Promotion of Science (21H02115)
Bio-SPMs Collaborative Research of the WPI Nano Life Science Institute
Japan Agency for Medical Research and Development (JP20am0101070 (support number 1841))
Article History
Received: 25 August 2025
Accepted: 11 May 2026
First Online: 15 June 2026
Competing interests
: The authors declare no competing interests.