Permana, Andrea L. https://orcid.org/0000-0002-8010-1439
Permana, Junaidi Jaka
Nellissen, Lara
Prayogi, Eggi Septian
Prasetyo, Didik
Wich, Serge A. https://orcid.org/0000-0003-3954-5174
van Schaik, Carel P.
Schuppli, Caroline https://orcid.org/0000-0001-6318-2815
Funding for this research was provided by:
Leakey Foundation
Universität Zürich
Article History
Received: 8 November 2024
Accepted: 13 May 2025
First Online: 7 June 2025
Competing interests
: The authors declare no competing interests.
: We the authors affirm that this research was conducted in accordance with the highest ethical standards in biological research. All applicable institutional and national guidelines for the study of wild animals were followed. Ethical approval for our research was granted by the Indonesian Institute of Science (LIPI), the Indonesian State Ministry for Research and Technology (RISTEKDIKTI), the National Research and Innovation Agency (BRIN), the Directorate General of Natural Resources and Ecosystem Conservation under the Ministry of Environment and Forestry of Indonesia (KSDAE-KLHK) and the Gunung Leuser National Park (TNGL). All authors are committed to fostering an inclusive, collaborative research environment. Contributions to this work were made regardless of gender, race, ethnicity, nationality, disability, sexual orientation, or background. All authors support equity, diversity, and inclusion in science and acknowledge the importance of promoting underrepresented voices in biological research. We also confirm that the data and materials supporting this publication are made openly available in accordance with journal policies, promoting transparency and reproducibility.