Yoshida, Yosuke https://orcid.org/0009-0001-2999-1675
Machiyama, Hiroaki https://orcid.org/0000-0002-5524-581X
Wakamatsu, Ei https://orcid.org/0000-0003-2115-8520
Saito, Kensho
Takeuchi, Arata https://orcid.org/0000-0003-0412-533X
Nishikawa, Tetsushi
Matsushima, Ryohei https://orcid.org/0000-0002-6191-9753
Nishi, Wataru https://orcid.org/0000-0001-9919-8050
Nishijima, Hitoshi
Kanno, Yoshihiko
Yokosuka, Tadashi https://orcid.org/0000-0002-3914-3995
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (JP25113725, JP15H01194, JP16H06501, JP17H03600, JP19K22545, JP20H03536, JP23K27466, JP23H04790)
MEXT | Japan Society for the Promotion of Science (JP18K06165, JP24K10438)
MEXT | JST | Precursory Research for Embryonic Science and Technology
Naito Foundation
Research support from the Center for Diversity at Tokyo Medical University
Article History
Received: 2 June 2025
Accepted: 11 December 2025
First Online: 8 January 2026
Competing interests
: The authors declare no competing interests.