Zhang, Hui https://orcid.org/0009-0001-0052-0191
Feng, Huili https://orcid.org/0000-0001-9838-9787
Qu, Xinjing
Yang, Mei https://orcid.org/0009-0004-1827-1498
Huang, Mengke https://orcid.org/0009-0003-9476-7578
Jacobs, Douglass F.
Guo, Jiahuan https://orcid.org/0000-0002-3440-6696
Funding for this research was provided by:
Natural Science Foundation of Hainan Province (325QN230)
Natural Science Foundation of Hainan Province (425RC700)
National Natural Science Foundation of China (32301403)
Article History
Received: 1 September 2025
Accepted: 23 January 2026
First Online: 2 February 2026
Competing interests
: The authors declare no competing interests.