Chang, Vincent
Lian, Isaac
Mahoney, Keira E.
Romano, Jeffrey
Afshari, Ali Reza https://orcid.org/0009-0002-7365-3646
Chen, Ryan J.
Chen, Xiangjun
Fineide, Fredrik https://orcid.org/0000-0003-3835-8772
Khan, Ayyad Zartasht
Utheim, Tor Paaske
Karlsson, Niclas G. https://orcid.org/0000-0002-3045-2628
Laurie, Gordon W. https://orcid.org/0000-0002-5311-2650
Malaker, Stacy A. https://orcid.org/0000-0003-2382-5067
Funding for this research was provided by:
U.S. Department of Health & Human Services | NIH | National Institute of General Medical Sciences (R35-GM147039)
Article History
Received: 25 November 2025
Accepted: 13 May 2026
First Online: 26 May 2026
Competing interests
: The authors declare the following competing financial interest(s): S.A.M. is a co-inventor on a Stanford patent related to the use of mucinases as research tools. G.W.L. is Cofounder and CSO of TearSolutions, Inc. and IsletRegen, LLC and inventor of UVA patents on lacritin. All other authors declare no competing interests.