Yamaguchi, Tomoko https://orcid.org/0000-0001-9482-8738
Minamino, Tohru
Kato, Takayuki https://orcid.org/0000-0002-8879-6685
Chevance, Fabienne F. V. https://orcid.org/0000-0003-2997-3466
Hughes, Kelly T. https://orcid.org/0000-0001-5344-0972
Namba, Keiichi https://orcid.org/0000-0003-2911-5875
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (JP25000013)
MEXT | Japan Society for the Promotion of Science (JSPS Research Fellowship for Young Scientists)
MEXT | Japan Society for the Promotion of Science (JP19H03182)
MEXT | Japan Society for the Promotion of Science (JP22H02573)
MEXT | Japan Society for the Promotion of Science (JP22K19274)
MEXT | Japan Society for the Promotion of Science (JP20H05532)
MEXT | Japan Society for the Promotion of Science (JP22H04844)
Japan Agency for Medical Research and Development (JP21am0101117)
Japan Agency for Medical Research and Development (JP22ama121003)
Japan Agency for Medical Research and Development (JP17pc0101020)
JEOL YOKOGUSHI Research Alliance Laboratories of The University of Osaka
Article History
Received: 2 December 2025
Accepted: 30 June 2026
First Online: 13 July 2026
Competing interests
: The authors declare no competing interests.