Guo, Yiqun
Zhang, Jingkun
Jiang, Liangyu
Yin, Huamei
Niu, Xiankai
Shen, Liang https://orcid.org/0000-0003-1792-0105
Guo, Yuna https://orcid.org/0009-0006-7828-0884
Li, Yuancheng https://orcid.org/0000-0002-9833-9306
Funding for this research was provided by:
National Natural Science Foundation of China (22304104)
National Natural Science Foundation of China (22106091)
Natural Science Foundation of Shandong Province (ZR2022QH260)
Natural Science Foundation of Shandong Province (ZR2025MS109)
Natural Science Foundation of Shandong Province (ZR2021QB126)
Taishan Scholar Project of Shandong Province (tsqn202408255)
Article History
Received: 12 October 2025
Accepted: 5 March 2026
First Online: 25 March 2026
Competing interests
: The authors declare no competing interests.