Ren, Qiuping
Xie, Shang-Ping https://orcid.org/0000-0002-3676-1325
Peng, Qihua https://orcid.org/0000-0003-4913-3499
Li, Yuanlong https://orcid.org/0000-0002-7239-5756
Wang, Fan https://orcid.org/0000-0001-5932-7567
Funding for this research was provided by:
National Natural Science Foundation of China (42306037)
Natural Science Foundation of Shandong Province (ZR2023QD058)
China Postdoctoral Science Foundation (2023M743526)
China Postdoctoral Science Foundation (2024T170936)
Shandong Province Postdoctoral Innovation Talent Plan
National Natural Science Foundation of the Laoshan Laboratory
Article History
Received: 27 August 2025
Accepted: 9 September 2025
First Online: 17 October 2025
Competing interests
: The authors declare no competing interests.