Yang, Bo
Zhang, Fan https://orcid.org/0000-0003-1608-7764
Uyeshima, Makoto https://orcid.org/0000-0001-5831-2005
Lin, Jian https://orcid.org/0000-0002-6831-2014
Lin, Wule
Yang, Jianfeng https://orcid.org/0000-0002-8218-8080
Zhang, Xubo https://orcid.org/0000-0002-4015-9169
Wang, Wei
Liao, Chen
Fang, Hui
Qiu, Gengen
Hu, Xiangyun
Funding for this research was provided by:
CAS | South China Sea Institute of Oceanology, Chinese Academy of Sciences (2022YFF0802800)
CAS | South China Sea Institute of Oceanology, Chinese Academy of Sciences (2024B1515020100)
University of Tokyo (18403005)
Article History
Received: 19 August 2025
Accepted: 22 December 2025
First Online: 8 January 2026
Competing interests
: The authors declare no competing interests.