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Du, Chenliang https://orcid.org/0009-0007-7303-899X

Zhang, Ke https://orcid.org/0000-0002-1841-9016

Lin, Qi https://orcid.org/0000-0003-4977-8326

Huang, Shixin https://orcid.org/0000-0003-1062-1807

Han, Yaoyao https://orcid.org/0000-0003-4150-7855

Shen, Ji https://orcid.org/0000-0001-9143-5648

Funding for this research was provided by:

National Natural Science Foundation of China (42230507)

National Natural Science Foundation of China (42361144717)

National Natural Science Foundation of China (42477487)

National Natural Science Foundation of China (42501040)

National Natural Science Foundation of China (42230507)

Science and Technology Planning Project of Nanjing Institute of Geography and Limnology, Chinese Academy of Sciences

Basic Research Program of Jiangsu Province (BK20240213)

China Postdoctoral Science Foundation (GZC20231095)

Second Tibetan Plateau Scientific Expedition and Research Program (STEP)

Text and Data Mining valid from 2026-05-20

Accepted Manuscript valid from 2026-05-20