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Authors
Civiletto, Gabriele https://orcid.org/0000-0002-9818-9942
Brunetti, Dario https://orcid.org/0000-0002-2740-9370
Lizzo, Giulia
Muller, Kamila
Jacot, Guillaume E. https://orcid.org/0000-0001-5821-2717
Daskalaki, Ioanna
Sizzano, Federico
Huh, Minji
Di Meo, Ivano https://orcid.org/0000-0002-4616-5623
Colombo, Maria Nicol
Sanchez-Garcia, José L.
Bétrisey, Bertrand J.
Zollinger, Alix
Lino, Patricia
Neal, Christopher
Egesipe, Anne-Laure
Richard, Joy
Chimen, Myriam
Hermant, Aurélie
Brinon, Benjamin
Texari, Lorane
Metairon, Sylviane https://orcid.org/0000-0003-2434-8733
Qureshi, Mohammed Adnan
Patel, Dhaval S.
Vanapalli, Siva A.
Malavolta, Marco https://orcid.org/0000-0002-8442-1763
Gao, Arwen W. https://orcid.org/0000-0001-7687-8926
Lalou, Amelia
Provinciali, Mauro
Orlando, Fiorenza
Tiranti, Valeria https://orcid.org/0000-0002-3584-7338
Brooke, Robert T.
Horvath, Steve https://orcid.org/0000-0002-4110-3589
Auwerx, Johan
Feige, Jerome N. https://orcid.org/0000-0002-4751-264X
Gut, Philipp https://orcid.org/0000-0002-0689-9518
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Article History
Received: 14 August 2023
Accepted: 8 August 2025
First Online: 24 September 2025
Competing interests
: G.C. is an employee of DSM-Firmenich. G.L., K.M., G.E.J., F.S., M.H., J.L.S.-G., B.J.B., P.L., B.B., A.-L.E., A.Z., J.R., A.H., L.T., S.M., M.C., J.N.F. and P.G. are employees of Nestlé Research, part of Société des Produits Nestlé SA. Patents owned by Société des Produits Nestlé SA have been filed related to this work, and the list of inventors includes authors of this article. Patent application publications can be found using the following identifiers: , and . M.A.Q., D.S.P. and S.A.V. are employees of NemaLife Inc., which has received fees from Nestlé Research for preclinical services. The Regents of the University of California are the sole owners of patents and patent applications directed at epigenetic biomarkers and the mammalian array for which S.H. is a named inventor. S.H. is a founder and paid consultant of the nonprofit Epigenetic Clock Development Foundation, which licenses these patents. No new patents were filed for epigenetic age analysis as part of this study. S.H. is a principal investigator at Altos Labs. R.T.B. is a cofounder and employee of the nonprofit Epigenetic Clock Development Foundation. J.A. is a consultant and/or scientific advisory board member of MitoBridge/Astellas, MetroBiotech, Amazentis, Vandria, OrsoBio and NovMetapharma. The other authors declare no competing interests.
