Kim, Minhoo https://orcid.org/0000-0002-4783-4884
Wang, Justin
Pilley, Steven E. https://orcid.org/0000-0001-9670-3314
Lu, Ryan J. https://orcid.org/0000-0001-8627-2880
Xu, Alan https://orcid.org/0000-0002-8698-9301
Kim, Younggyun
Liu, Minying
Fu, Xueyan
Booth, Sarah L.
Mullen, Peter J.
Benayoun, Bérénice A. https://orcid.org/0000-0002-7401-4777
Funding for this research was provided by:
Pew Charitable Trusts (#00034120)
GCRLE: GCRLE-2020
USC Provost's Undergraduate Research Fellowship
U.S. Department of Health & Human Services | NIH | National Institute on Aging (T32 AG052374)
Diana Jacobs Kalman/AFAR Scholarships for Research in the Biology of Aging
United States Department of Agriculture | Agricultural Research Service (No. 58-1950-7-707)
Article History
Received: 10 May 2024
Accepted: 7 January 2026
First Online: 3 March 2026
Competing interests
: The authors declare no competing interests.