Hori, Tetsuya https://orcid.org/0000-0002-0363-4688
Mahana, Yutaka https://orcid.org/0000-0001-6218-6726
Ariyoshi, Mariko https://orcid.org/0000-0002-1361-2642
Fukagawa, Tatsuo https://orcid.org/0000-0001-8564-6852
Funding for this research was provided by:
MEXT | JST | Core Research for Evolutional Science and Technology of Japan Science and Technology Agency (JPMJCR21E6)
MEXT | Japan Society for the Promotion of Science (22H00408)
MEXT | Japan Society for the Promotion of Science (23K18113)
MEXT | Japan Society for the Promotion of Science (24H02281)
MEXT | Japan Society for the Promotion of Science (25H00975)
MEXT | Japan Society for the Promotion of Science (23K27153)
MEXT | Japan Society for the Promotion of Science (24K23185)
MEXT | Japan Society for the Promotion of Science (24K09340)
Article History
Received: 25 May 2025
Revised: 18 November 2025
Accepted: 8 December 2025
First Online: 8 January 2026
Disclosure and competing interests statement
: The authors declare no competing interests.