Fu, Hui-Hui https://orcid.org/0000-0001-8101-863X
Wang, Ming-Chen https://orcid.org/0009-0009-2071-5672
Wang, Zhi-Qing
Sang, Yu-Han
Li, Zhen-Kun
Li, Fei-Fei
Liu, Jia-Rong
Qin, Qi-Long
Zhu, Xiao-Yu
Wang, Na
Wan, Jin-Jian
Teng, Zhao-Jie
Zhang, Wei-Peng
Gates, Andrew J https://orcid.org/0000-0002-4594-5038
Li, Chun-Yang https://orcid.org/0000-0002-1151-4897
Todd, Jonathan D https://orcid.org/0000-0003-0777-9312
Zhang, Yu-Zhong https://orcid.org/0000-0002-2017-1005
Funding for this research was provided by:
National Key R & D program of China (2022YFC2807501)
National Key R & D program of China (2024YFC2816000)
National Science Foundation of China (W2441012)
National Science Foundation of China (32330001)
National Science Foundation of China (92251303)
National Science Foundation of China (42176156)
National Science Foundation of China (42276102)
Fundamental Research Funds for the Central Universities (202172002)
Major Scientific and Technological Innovation Project (MSTIP) of Shandong Province (2019JZZY010817)
Taishan Scholars Program of Shandong Province, China (tspd20240806)
Taishan Scholars Program of Shandong Province, China (tsqn202306092)
National Science Foundation of Shandong Province (ZR2025MS308)
National Science Foundation of Shandong Province (ZR2024YQ076)
Biotechnology and Biological Sciences Research Council, UK (BB/X005968)
Natural Environmental Research Council, UK (NE/P012671)
Natural Environmental Research Council, UK (NE/S001352)
Natural Environmental Research Council, UK (NE/X000990)
Natural Environmental Research Council, UK (NE/X014428 with DLS)
the Leverhulme trust grants (RPG-2020-413)
Article History
Received: 15 August 2025
Revised: 19 December 2025
Accepted: 9 January 2026
First Online: 9 February 2026
Disclosure and competing interests statement
: The authors declare no competing interests.