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Xie, Guohui https://orcid.org/0000-0003-4734-9114

Du, Xuan https://orcid.org/0000-0001-9589-3566

Tan, Yifang

Zhou, Yuxing https://orcid.org/0000-0002-9065-7908

Chi, Cheng

Zhou, Sixian

Picard, Colette L https://orcid.org/0000-0002-2177-2216

Chai, Songge https://orcid.org/0009-0001-6773-8005

Wu, Lei https://orcid.org/0009-0003-9757-3795

Zhu, Danling https://orcid.org/0000-0003-3385-403X

Zhao, Jun https://orcid.org/0009-0006-6190-0469

Xue, Yan https://orcid.org/0000-0003-0871-8853

Li, Sisi https://orcid.org/0000-0002-7290-8128

Jacobsen, Steven E https://orcid.org/0000-0001-9483-138X

Wu, Zhe https://orcid.org/0000-0001-9436-3299

Du, Jiamu https://orcid.org/0000-0002-1337-0786

Funding for this research was provided by:

MOST | National Natural Science Foundation of China (32325008)

MOST | National Natural Science Foundation of China (323B2041)

GDSTC | Natural Science Foundation of Guangdong Province (2023B1515120048)

中国博士后科学基金会职 | Postdoctoral Research Foundation of China (2024M761288)

Shenzhen Medical Research Fund (A2303024)

深圳市科技创新委员会 | Natural Science Foundation of Shenzhen Municipality (RCJC20221008092720004)

深圳市科技创新委员会 | Natural Science Foundation of Shenzhen Municipality (RCJC20231211085945061)

深圳市科技创新委员会 | Natural Science Foundation of Shenzhen Municipality (RCJC20231211085924032)

深圳市科技创新委员会 | Natural Science Foundation of Shenzhen Municipality (JCYJ20240813094610015)

深圳市科技创新委员会 | Natural Science Foundation of Shenzhen Municipality (ZDSYS20230626091659010)

SUSTech Institute for Biological Electron Microscopy (N.A.)

Howard Hughes Medical Institute (N.A.)

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Version of Record valid from 2026-04-02