Komatsu-Hirota, Satoko
Tabata, Keisuke
Sou, Yu-shin
Kakuta, Soichiro https://orcid.org/0000-0001-9434-4944
Sakamaki, Jun-ichi https://orcid.org/0000-0003-3361-6314
Tsuchiya, Hikaru https://orcid.org/0000-0002-2797-6711
Li, Jiachen https://orcid.org/0009-0005-4666-6486
Kumeta, Hiroyuki https://orcid.org/0000-0003-3713-2122
Sakai, Yuji
Fujioka, Yuko https://orcid.org/0000-0002-6905-0669
Noshiro, Daisuke https://orcid.org/0000-0001-7024-2054
Shimobayashi, Shunsuke F
Kurimura, Tomo https://orcid.org/0009-0005-7858-6413
Taniguchi, Takashi https://orcid.org/0000-0003-0017-7669
Abe, Manabu
Koike, Masato https://orcid.org/0000-0002-3174-5684
Morishita, Hideaki
Noda, Nobuo N https://orcid.org/0000-0002-6940-8069
Komatsu, Masaaki https://orcid.org/0000-0001-7672-7722
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (JP23K20044)
MEXT | Japan Society for the Promotion of Science (JP24H00060)
MEXT | Japan Society for the Promotion of Science (JP25H01323)
MEXT | Japan Society for the Promotion of Science (JP24H01901)
MEXT | Japan Society for the Promotion of Science (JP23K27134)
MEXT | Japan Society for the Promotion of Science (JP23H04923)
MEXT | Japan Society for the Promotion of Science (JP25K09567)
MEXT | Japan Society for the Promotion of Science (JP23K06065)
MEXT | Japan Society for the Promotion of Science (JP25H01419)
MEXT | Japan Society for the Promotion of Science (JP25H00966)
MEXT | Japan Society for the Promotion of Science (JP25H01320)
MEXT | Japan Society for the Promotion of Science (JP25H0132)
MEXT | Japan Society for the Promotion of Science (JP22H04926)
MEXT | Japan Society for the Promotion of Science (JP25H00315)
MEXT | Japan Society for the Promotion of Science (JP25H01328)
Japan Agency for Medical Research and Development (JP22gm1410004h0003)
Japan Agency for Medical Research and Development (21gm6410019h0001)
MEXT | JST | Core Research for Evolutional Science and Technology (JPMJCR20E3)
Takeda Medical Research Foundation
Uehara Memorial Foundation
Kobayashi Foundation for Cancer Research
Mitsubishi Foundation
Article History
Received: 22 December 2025
Revised: 24 March 2026
Accepted: 2 April 2026
First Online: 5 May 2026
Disclosure and competing interests statement
: The authors declare no competing interests.