Zheng, Yu https://orcid.org/0009-0001-5466-7762
Chai, Simin
Zhong, Cuijuan https://orcid.org/0009-0003-2601-8620
Sun, Yixuan
Xu, Shixia
Ren, Wenhua
Yang, Guang https://orcid.org/0000-0001-6285-6937
Funding for this research was provided by:
MOST | National Natural Science Foundation of China (32030011)
MOST | National Natural Science Foundation of China (32270442)
MOST | National Natural Science Foundation of China (32070409)
MOST | National Natural Science Foundation of China (U24A20362)
MOST | National Key Research and Development Program of China (2022YFF1301600)
Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou) (GML2021 GD0805)
Priority Academic Program Development of Jiangsu Higher Education Institutions
Article History
Received: 9 March 2025
Revised: 18 October 2025
Accepted: 28 October 2025
First Online: 11 November 2025
Disclosure and competing interests statement
: The authors declare no competing interests.