Furukawa, Kentaro https://orcid.org/0000-0002-1551-1538
Maruyama, Tatsuro https://orcid.org/0009-0007-2550-9064
Sakai, Yuji https://orcid.org/0000-0002-2184-4304
Yamashita, Shun-ichi https://orcid.org/0000-0001-8892-0765
Inoue, Keiichi https://orcid.org/0000-0002-7101-486X
Fukuda, Tomoyuki https://orcid.org/0000-0003-2069-7127
Noda, Nobuo N https://orcid.org/0000-0002-6940-8069
Kanki, Tomotake https://orcid.org/0000-0001-9646-5379
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (JP23K20044,JP24H00060,JP25H00966,JP25H01321,JP25H01320)
MEXT | Japan Society for the Promotion of Science (JP23K23878,JP24H02274)
MEXT | Japan Society for the Promotion of Science (JP25K21747)
MEXT | Japan Society for the Promotion of Science (JP25H01101)
MEXT | Japan Society for the Promotion of Science (JP23H04255)
MEXT | Japan Society for the Promotion of Science (JP24K01717)
MEXT | Japan Society for the Promotion of Science (JP24K09373)
MEXT | Japan Society for the Promotion of Science (JP23K05715)
MEXT | Japan Society for the Promotion of Science (JP24K01980)
MEXT | Japan Society for the Promotion of Science (JP25K09559)
Japan Agency for Medical Research and Development (JP23gm1710006s0101)
MEXT | Japan Science and Technology Agency (JPMJCR20E3)
Takeda Science Foundation
Institute for Fermentation, Osaka
Noda Institute for Scientific Research
Article History
Received: 16 June 2025
Revised: 8 December 2025
Accepted: 15 December 2025
First Online: 13 January 2026
Disclosure and competing interests statement
: The authors declare no competing interests.