Tsuji, Shunya https://orcid.org/0009-0004-3169-4647
Nakano, Sosuke
Ito, Koyu
Minami, Shohei
Uemura, Ken
Konishi, Yusuke
Wakita, Masahiro
Okumura, Yumiko
Kawamoto, Shimpei https://orcid.org/0000-0002-7033-671X
Matsuki, Akari https://orcid.org/0009-0003-8786-9189
Nakaoka, Shinji
Ono, Chikako https://orcid.org/0000-0002-3965-6715
Takahashi, Hiroo
Anzai, Itsuki https://orcid.org/0000-0001-9307-943X
Watanabe, Tokiko
Uezumi, Akiyoshi
Matsuura, Yoshiharu
Kobayashi, Takeshi
Okamoto, Toru https://orcid.org/0000-0003-4000-3102
Tsuboi, Akio https://orcid.org/0000-0002-4790-4512
Asagiri, Masataka
Hara, Eiji https://orcid.org/0000-0001-7821-3960
Funding for this research was provided by:
Mitsubishi Foundation (202012016)
Japan Agency for Medical Research and Development (JP25zf0127008h0002)
Japan Agency for Medical Research and Development (JP25gm1710004h0004)
MEXT | Japan Science and Technology Agency (JPMJMS2022)
MEXT | Japan Science and Technology Agency (JPMJFR2308)
MEXT | Japan Society for the Promotion of Science (JP25H00443)
MEXT | Japan Society for the Promotion of Science (JP23K06481)
MEXT | Japan Society for the Promotion of Science (JP25K00099)
Takeda Science Foundation
Uehara Memorial Foundation
Lotte Foundation
Ministry of Education, Culture, Sports, Science and Technology (JPMXP1323015484)
Nagase Science and Technology Foundation
OU Master Plan Implementation Project promoted under the University of Osaka
Article History
Received: 23 January 2026
Revised: 24 March 2026
Accepted: 26 March 2026
First Online: 10 April 2026
Disclosure and competing interests statement
: The authors declare no competing interests.