Moon, Young Jae https://orcid.org/0000-0002-5285-7618
McCormack, Nikki https://orcid.org/0000-0002-7454-3732
Diehn, Thomas https://orcid.org/0009-0008-8018-7819
Lin, Stephen
Staedler, Katia
Bhattacharya, Surajit
Bittel, Daniel C
Wang, Paul C https://orcid.org/0000-0001-8219-7051
Hogarth, Marshall
Oh, Jong-Sung
Goh, Tae Sik
Min, Ji Hong
Stojkovic, Tanya https://orcid.org/0000-0002-4054-2838
Ackermann-Bonan, Isabelle
Jaiswal, Jyoti K https://orcid.org/0000-0002-5992-5185
Funding for this research was provided by:
HHS | NIH | National Institute of Arthritis and Musculoskeletal and Skin Diseases (R01AR055686)
HHS | NIH | National Institute of Arthritis and Musculoskeletal and Skin Diseases (K01AR077686)
HHS | NIH | Eunice Kennedy Shriver National Institute of Child Health and Human Development (P50HD105328)
Ministry of Science and ICT, South Korea (2022R1C1C1006721)
Jeonbuk National University (54896)
Biomedical Research Institute, Jeonbuk National University Hospital (54907)
Article History
Received: 9 October 2025
Revised: 20 August 2026
Accepted: 21 August 2026
First Online: 12 October 2026
Disclosure and competing interests statement
: The authors JKJ and MWH hold US Patent #12,594,278, which is related to the work reported here. The remaining authors declare no competing interests.