Rutz, Christian
Klump, Barbara C.
Komarczyk, Lisa
Leighton, Rosanna
Kramer, Joshua
Wischnewski, Saskia
Sugasawa, Shoko
Morrissey, Michael B.
James, Richard
St Clair, James J. H.
Switzer, Richard A.
Masuda, Bryce M.
Article History
Received: 16 February 2016
Accepted: 12 July 2016
First Online: 14 September 2016
Competing interests
: The authors declare the following competing financial interests: five co-authors were (R.L., J.K., R.A.S.), or still are (L.K., B.M.M.), employees of San Diego Zoo Global, which is a not-for-profit organisation.