Updates are available Erratum dated 2017-02-01

Article History

Received: 18 August 2015

Accepted: 15 October 2015

First Online: 17 December 2015

Change Date: 24 October 2016

Change Type: Corrigendum

Change Details: In the version of this article initially published, the citation of Figure 3 in the second paragraph of the third subsection (Chromatin: the nexus of phenotype and the environment) is incorrect. That should cite Figure 2, as follows: "From an evolutionary perspective, two extreme models of how such complexity might be generated and regulated can be envisaged95(Fig. 2)." The error has been corrected in the HTML and PDF versions of the article.

Competing interests

: The authors declare no competing financial interests.