Received: 12 June 2015
Accepted: 8 October 2015
First Online: 9 November 2015
Change Date: 30 March 2020
Change Type: Clarification
Change Date: 20 November 2015
Change Type: Corrigendum
Change Details: Editors’ note, March 2020: We are aware that this article is being used as the basis for unverified theories that the novel coronavirus causing COVID-19 was engineered. There is no evidence that this is true; scientists believe that an animal is the most likely source of the coronavirus.
Change Details: In the version of this article initially published online, the authors omitted to acknowledge a funding source, USAID-EPT-PREDICT funding from EcoHealth Alliance, to Z.-L.S. The error has been corrected for the print, PDF and HTML versions of this article.
: The authors declare no competing financial interests.
Free to read: This content has been made available to all.