Article History
Received: 23 May 2014
Accepted: 12 September 2014
First Online: 12 October 2014
Change Date: 10 August 2015
Change Type: Correction
Change Details: In the version of this article initially published, a minus sign was missing before the coefficient <i>β</i> in equation (3), Online Methods. The error has been corrected in the HTML and PDF versions of the article.
Competing interests
: The authors declare no competing financial interests.