Funding for this research was provided by:
Natural Environment Research Council (NERC) (NE/J008745/1)
Natural Environment Research Council(UK)
Received: 20 April 2017
Accepted: 3 July 2018
First Online: 7 August 2018
Change Date: 5 September 2018
Change Type: Update
Change Details: This Article was originally published without the accompanying Peer Review File. This file is now available in the HTML version of the Article; the PDF was correct from the time of publication.
: The authors declare no competing interests.