Updates are available Correction dated 2020-07-17
Click to view Correction: https://doi.org/10.1038/s41586-020-2556-6
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Authors
Thaventhiran, James E. D. http://orcid.org/0000-0001-8616-074X
Lango Allen, Hana http://orcid.org/0000-0002-7803-8688
Burren, Oliver S.
Rae, William
Greene, Daniel
Staples, Emily
Zhang, Zinan
Farmery, James H. R.
Simeoni, Ilenia
Rivers, Elizabeth
Maimaris, Jesmeen
Penkett, Christopher J. http://orcid.org/0000-0003-4006-7261
Stephens, Jonathan
Deevi, Sri V. V. http://orcid.org/0000-0002-0405-4335
Sanchis-Juan, Alba
Gleadall, Nicholas S.
Thomas, Moira J.
Sargur, Ravishankar B.
Gordins, Pavels
Baxendale, Helen E.
Brown, Matthew
Tuijnenburg, Paul
Worth, Austen
Hanson, Steven
Linger, Rachel J.
Buckland, Matthew S. http://orcid.org/0000-0002-5646-4707
Rayner-Matthews, Paula J.
Gilmour, Kimberly C.
Samarghitean, Crina
Seneviratne, Suranjith L.
Sansom, David M.
Lynch, Andy G. http://orcid.org/0000-0002-7876-7338
Megy, Karyn http://orcid.org/0000-0002-2826-3879
Ellinghaus, Eva http://orcid.org/0000-0003-2914-3382
Ellinghaus, David http://orcid.org/0000-0002-4332-6110
Jorgensen, Silje F.
Karlsen, Tom H.
Stirrups, Kathleen E. http://orcid.org/0000-0002-6823-3252
Cutler, Antony J.
Kumararatne, Dinakantha S.
Chandra, Anita
Edgar, J. David M.
Herwadkar, Archana
Cooper, Nichola
Grigoriadou, Sofia
Huissoon, Aarnoud P.
Goddard, Sarah
Jolles, Stephen
Schuetz, Catharina
Boschann, Felix http://orcid.org/0000-0001-9410-9290
Abbs, Stephen
Adhya, Zoe
Adlard, Julian
Afzal, Maryam
Ahmed, Irshad
Ahmed, Munaza
Ahmed, Saeed
Aitman, Timothy J.
Alachkar, Hana
Alamelu, Jayanthi
Alikhan, Raza
Allen, Carl E.
Allen, Louise
Allsup, David J.
Alvi, Arif
Ambegaonkar, Gautam
Anantharachagan, Ariharan
Ancliff, Philip
Anderson, Julie
Antrobus, Richard
Armstrong, Ruth
Arno, Gavin
Arumugakani, Gururaj
Arya, Rita
Ashford, Sofie
Astle, William J.
Attwood, Anthony
Austin, Steve
Aydinok, Yesim
Ayub, Waqar
Babbs, Christian
Bacchelli, Chiara
Baglin, Trevor
Bakchoul, Tamam
Bariana, Tadbir K.
Barratt, Jonathan
Barwell, Julian
Baski, John
Bates, Rachel W.
Batista, Joana
Baxendale, Helen E.
Baynam, Gareth
Bennett, David L.
Bethune, Claire
Bhatnagar, Neha
Bibi, Shahnaz
Bierzynska, Agnieszka
Biss, Tina
Bitner-Glindzicz, Maria A. K.
Bleda, Marta
Blesneac, Iulia
Boardman, Barbara
Boddana, Preetham
Bogaard, Harm J.
Booth, Claire
Boyce, Sara
Bradley, John R.
Brady, Angela
Breen, Gerome
Brennan, Paul
Brewer, Carole
Briley, Annette
Brown, Matthew
Brown, Richard
Browning, Michael J.
Brownlie, Mary
Bryson, Christine J.
Buchan, Rachel J.
Buck, Jackie
Buckland, Matthew S.
Bueser, Teofila
Diz, Carmen Bugarin
Burns, Siobhan O.
Burren, Oliver S.
Calleja, Paul
Carmichael, Jenny
Carr-White, Gerald
Carss, Keren J.
Casey, Ruth
Chalmers, Elizabeth
Chambers, Jenny
Chambers, John
Chan, Melanie M. Y.
Chan, Melissa V.
Chandra, Anita
Cheng, Floria
Chinn, Ivan K.
Chinnery, Patrick F.
Chitre, Manali
Chong, Sam
Christian, Martin T.
Church, Colin
Clement, Emma M.
Brod, Naomi Clements
Clifford, Hayley
Clowes, Virginia E.
Coghlan, Gerry
Colby, Elizabeth
Cole, Trevor R. P.
Collins, Janine H.
Collins, Peter W.
Condliffe, Robin
Cook, H. Terence.
Cook, Stuart
Cookson, Victoria
Cooper, Nichola
Corris, Paul A.
Creaser-Myers, Amanda
Crisp-Hihn, Abigail
Curry, Nicola S.
Cutler, Antony J.
Da Costa, Rosa
Danesino, Cesare
Daniels, Matthew J.
Darby, Damaris
Daugherty, Louise C.
Davies, E. G.
Davies, Sophie
Davis, John
de Bree, Godelieve J.
Deacock, Sarah
Deegan, Patrick B.
Deevi, Sri V. V.
Dempster, John
Dent, Timothy
Deshpande, Charu
Devlin, Lisa A.
Dewhurst, Eleanor F.
Dixit, Anand K.
Dixon, Peter H.
Doffinger, Rainer
Dolling, Helen
Dormand, Natalie
Downes, Kate
Drazyk, Anna M.
Drewe, Elizabeth
Duarte, Daniel
Dutt, Tina
Edgar, J. David M.
Edwards, Karen E.
Egner, William
Ekani, Melanie N.
El-Shanawany, Tariq
Elkhalifa, Shuayb
Elston, Tony
Emmerson, Ingrid
Erber, Wendy N.
Erwood, Marie
Estiu, Maria C.
Evans, Dafydd Gareth
Evans, Gillian
Everington, Tamara
Eyries, Mélanie
Farmery, James H. R.
Favier, Remi
Firth, Helen V.
Fitzpatrick, Maggie M.
Fletcher, Debra
Flinter, Frances A.
Fox, James C.
Frary, Amy J.
French, Courtney E.
Freson, Kathleen
Frontini, Mattia
Furie, Bruce
Gale, Daniel P.
Gall, Henning J.
Gardham, Alice
Gaspar, H. Bobby
Gattens, Michael
Ghali, Neeti
Ghataorhe, Pavandeep K.
Ghio, Stefano
Ghofrani, Hossein-Ardeschir
Ghurye, Rohit
Gibbs, J. Simon R.
Gilbert, Rodney D.
Gilmour, Kimberly C.
Girerd, Barbara
Girling, Joanna C.
Gissen, Paul
Gleadall, Nicholas S.
Goddard, Sarah
Gordins, Pavels
Gorman, Kathleen M.
Gosal, David
Graf, Stefan
Grassi, Luigi
Greene, Daniel
Greenhalgh, Alan J.
Greenhalgh, Lynn
Greinacher, Andreas
Gresele, Paolo
Griffiths, Philip G.
Griffiths, Sian
Grigoriadou, Sofia
Grozeva, Detelina
Hackett, Scott J.
Hadden, Robert D. M.
Hadinnapola, Charaka
Hague, Rosie
Hague, William M.
Haimel, Matthias
Hall, Matthew
Halmagyi, Csaba
Hammerton, Tracey
Hanson, Helen L.
Harkness, Kirsty
Harper, Andrew R.
Harper, Lorraine
Harris, Claire
Harrison, Claire
Hart, Daniel
Hassan, Ahamad
Hayman, Grant
Heemskerk, Johan W. M.
Hegde, Shivaram
Henderson, Alex
Henderson, Robert H.
Hensiek, Anke
Henskens, Yvonne M. C.
Herwadkar, Archana
Hodgson, Joshua
Hoffman, Jonathan
Holden, Simon
Holder, Muriel
Horvath, Rita
Houlden, Henry
Houweling, Arjan C.
Howard, Luke S.
Hu, Fengyuan
Hudson, Gavin
Hughes, Sean
Hughes, Stephen
Huis in ‘t Veld, Anna E.
Huissoon, Aarnoud P.
Humbert, Marc
Hurles, Matthew E.
Hurst, Jane A.
Irvine, Val
Izatt, Louise
James, Roger
Jeevaratnam, Praveen
Johnson, Mark
Johnson, Sally A.
Jolles, Stephen
Jolley, Jennifer D.
Jones, Bryony
Jones, Julie
Josifova, Dragana
Jurkute, Neringa
Karim, Yousuf M.
Karoshi, Mahantesh A.
Kasanicki, Mary A.
Kazkaz, Hanadi
Kazmi, Rashid
Keeling, David
Kelleher, Peter
Kelly, Anne M.
Kempster, Carly
Kennedy, Fiona
Kiani, Sorena
Kiely, David G.
Kingston, Nathalie
Kinsey, Sally
Klein, Nigel
Klima, Robert
Knox, Ellen
Kostadima, Myrto A.
Kovacs, Gabor
Koziell, Ania B.
Kreuzhuber, Roman
Krishnakumar, Deepa
Kuijpers, Taco W.
Kumar, Ajith
Kumararatne, Dinakantha S.
Kurian, Manju A.
Laffan, James
Laffan, Michael A.
Lalloo, Fiona
Lambert, Michele P.
Lango Allen, Hana
Lawman, Sarah H. A.
Lawrie, Allan
Layton, D. Mark
Lear, Sara E.
Lees, Melissa M.
Lentaigne, Claire
Levine, Adam P.
Lewington, Andrew J. P.
Li, Wei
Liesner, Ri
Linger, Rachel J.
Liu, Bin
Longhurst, Hilary
Lorenzo, Lorena E.
Louka, Eleni
Hadeler, Silvia Lucato
Lyons, Paul A.
Macdougall, Malcolm
Machado, Rajiv D.
MacKenzie Ross, Robert V.
Mackillop, Lucy H.
MacLaren, Robert
Madan, Bella
Magee, Laura
Mahdi-Rogers, Mohamed
Maher, Eamonn R.
Maimaris, Jesmeen
Makris, Mike
Mangles, Sarah
Manson, Ania
Manzur, Adnan
Mapeta, Rutendo
Marchbank, Kevin J.
Mark, Patrick B.
Marks, Stephen
Markus, Hugh S.
Marschall, Hanns-Ulrich
Marshall, Andrew
Martin, Jennifer M.
Masati, Larahmie
Mathias, Mary
Matser, Vera
Matthews, Emma L.
Maw, Anna
Maxwell, Heather
McAlinden, Paul
McCarthy, Mark I.
McDermott, Elizabeth M.
McGowan, Simon J.
McJannet, Coleen
McKinney, Harriet
Meacham, Stuart
Mead, Adam J.
Castello, Ignacio Medina
Meehan, Sharon
Megy, Karyn
Mehta, Sarju
Mercer, Catherine L.
Michaelides, Michel
Michell, Anna C.
Milford, David
Millar, Carolyn M.
Millar, Hazel
Mistry, Anoop
Moenen, Floor
Moledina, Shahin
Montani, David
Moore, Anthony T.
Moore, Jason
Morrell, Nicholas W.
Morrisson, Valerie
Mozere, Monika
Muir, Keith W.
Mumford, Andrew D.
Murng, Sai H. K.
Nasir, Iman
Nejentsev, Sergey
Newnham, Michael
Ng, Joanne
Ngoh, Adeline
Noorani, Sadia
Noori, Muna
Nurden, Paquita
O’Sullivan, Jennifer M.
Obaji, Samya
Okoli, Steven
Oksenhendler, Eric
Olschewski, Andrea
Olschewski, Horst
Ong, Albert C. M.
Ong, Kai Ren
Oram, Helen
Ormondroyd, Elizabeth
Othman, Shokri
Ouwehand, Willem H.
Pantazis, Antonis
Papadia, Sofia
Papandreou, Apostolos
Park, Soo-Mi
Parker, Alasdair P. J.
Parry, David
Parsons, Georgina
Pasi, K. John.
Paterson, Joan
Payne, Jeanette H.
Peacock, Andrew J.
Peerlinck, Kathelijne
Penkett, Christopher J.
Pepke-Zaba, Joanna
Perry, David
Petersen, Romina
Piechowski-Jozwiak, Bartlomiej
Pinto, Fernando
Polwarth, Gary J.
Ponsford, Mark J.
Prasad, Sanjay
Prokopenko, Inga
Psaila, Beth
Pyle, Angela
Qasim, Waseem
Quinn, Ellen
Quinti, Isabella
Raina, Sanjay
Ranganathan, Lavanya
Rankin, Julia
Rankin, Stuart
Rao, Anupama
Raymond, F. Lucy
Rayner-Matthews, Paula J.
Rehnstrom, Karola
Reid, Evan
Reilly, Mary M.
Renton, Tara
Revel-Vilk, Shoshana
Rhodes, Christopher J.
Rice, Andrew S. C.
Richards, Emma E.
Richards, Mike
Richardson, Sylvia
Richter, Alex
Robert, Leema
Roberts, Irene
Rondina, Matthew T.
Rosser, Elisabeth
Rothwell, Peter
Roughley, Catherine
Roy, Noemi B.
Rue-Albrecht, Kevin
Sadeghi-Alavijeh, Omid
Saleem, Moin A.
Salmon, Richard M.
Samani, Nilesh J.
Samarghitean, Crina
Sambrook, Jennifer G.
Sanchis-Juan, Alba
Sandford, Richard
Santra, Saikat
Sargur, Ravishankar B.
Satchell, Simon C.
Savic, Sinisa
Scelsi, Laura
Schotte, Gwen
Schulman, Sol
Schulze, Harald
Scott, Richard
Scully, Marie
Searle, Claire
Seeger, Werner
Seneviratne, Suranjith L.
Sewell, W. A. Carrock
Seyres, Denis
Shackley, Fiona
Shamardina, Olga
Shapiro, Susan E.
Sharma, Pankaj
Shehata, Hassan A.
Shipley, Deborah
Shtoyerman, Rakefet
Sibson, Keith
Side, Lucy
Simeoni, Ilenia
Simpson, Michael
Sims, Matthew C.
Sinha, Manish D.
Sivapalaratnam, Suthesh
Skytte, Anne-Bine
Smith, Kenneth G. C.
Snape, Katie
Sneddon, Linda
Sohal, Aman
Soubrier, Florent
Southgate, Laura
Southwood, Mark
Splitt, Miranda
Staines, Simon
Staples, Emily
Stark, Hannah
Stauss, Hans
Steele, Cathal L.
Stein, Daniel
Stein, Penelope E.
Stephens, Jonathan
Stirrups, Kathleen E.
Stock, Sophie
Stubbs, Matthew J.
Suntharalingam, Jay
Swietlik, Emilia M.
Symington, Emily
Tait, R. Campbell
Talks, Kate
Tan, Rhea Y. Y.
Taylor, Gordon B.
Thachil, Jecko
Thaventhiran, James E. D.
Themistocleous, Andreas C.
Thomas, David C.
Thomas, Ellen
Thomas, Moira J.
Thomas, Patrick
Thompson, Dorothy A.
Thomson, Kate
Thrasher, Adrian J.
Thys, Chantal
Tilly, Tobias
Tischkowitz, Marc
Titterton, Catherine
Todd, John A.
Toh, Cheng-Hock
Tool, Anton T. J.
Toshner, Mark R.
Traylor, Matthew
Treacy, Carmen M.
Treadaway, Paul
Trembath, Richard C.
Trippier, Sarah
Tuna, Salih
Turek, Wojciech
Turro, Ernest
Upton, Paul D.
Urniaz, Rafal
Vale, Tom
Van Geet, Chris
van Zuydam, Natalie
Vandersteen, Anthony M.
Vazquez-Lopez, Marta
Veltman, Marijcke W. M.
Vogt, Julie
von Ziegenweidt, Julie
Vonk Noordegraaf, Anton
Vora, Ajay
Vries, Minka J. A.
Wakeling, Emma L.
Walker, Neil
Walker, Suellen M.
Walsh, Roddy
Wanjiku, Ivy
Ware, James S.
Warner, Timothy Q.
Wassmer, Evangeline
Watkins, Hugh
Watson, Henry G.
Watt, Christopher
Waugh, Dean
Webb, Nick
Webster, Andrew R.
Wei, Wei
Welch, Angela
Welch, Steven B.
Werring, David
Wessels, Julie
Westbury, Sarah K.
Westwood, John-Paul W.
Wharton, John
Whitehorn, Deborah
Whitworth, James
Wilkins, Martin R.
Willcocks, Lisa
Williams, David J.
Williamson, Catherine
Wong, Edwin K. S.
Wood, Nicholas
Wood, Yvette
Woods, Christopher Geoffrey
Woodward, Emma R.
Workman, Sarita
Wort, Stephen J.
Worth, Austen
Yates, Katherine
Yeatman, Nigel
Yong, Patrick F. K.
Young, Timothy
Yu, Ping
Yu-Wai-Man, Patrick
Zlamalova, Eliska
Lyons, Paul A. http://orcid.org/0000-0001-7035-8997
Hurles, Matthew E.
Savic, Sinisa http://orcid.org/0000-0001-7910-0554
Burns, Siobhan O.
Kuijpers, Taco W.
Turro, Ernest http://orcid.org/0000-0002-1820-6563
Ouwehand, Willem H. http://orcid.org/0000-0002-7744-1790
Thrasher, Adrian J. http://orcid.org/0000-0002-6097-6115
Smith, Kenneth G. C.
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Article History
Received: 1 December 2018
Accepted: 26 February 2020
First Online: 6 May 2020
Change Date: 17 July 2020
Change Type: Correction
Change Details: A Correction to this paper has been published:
Change Details: https://doi.org/10.1038/s41586-020-2556-6
Competing interests
: The authors declare no competing interests.
