Funding for this research was provided by:
Wellcome Trust (203141/Z/16/Z, 212284/Z/18/Z, 098387/Z/12/Z)
Engineering and Physical Sciences Research Council (EP/G03706X/1)
Received: 20 February 2019
Accepted: 15 July 2019
First Online: 2 September 2019
: S.R.M. is a director of GENSCI limited. The remaining authors declare no competing financial interests.