Moqri, Mahdi http://orcid.org/0000-0001-6675-0566
Herzog, Chiara http://orcid.org/0000-0002-1572-498X
Poganik, Jesse R. http://orcid.org/0000-0003-3098-8550
Ying, Kejun http://orcid.org/0000-0002-1791-6176
Justice, Jamie N.
Belsky, Daniel W. http://orcid.org/0000-0001-5463-2212
Higgins-Chen, Albert T. http://orcid.org/0000-0003-2904-2741
Chen, Brian H.
Cohen, Alan A. http://orcid.org/0000-0003-4113-3988
Fuellen, Georg
Hägg, Sara http://orcid.org/0000-0002-2452-1500
Marioni, Riccardo E. http://orcid.org/0000-0003-4430-4260
Widschwendter, Martin http://orcid.org/0000-0002-7778-8380
Fortney, Kristen
Fedichev, Peter O.
Zhavoronkov, Alex
Barzilai, Nir
Lasky-Su, Jessica http://orcid.org/0000-0001-6236-4705
Kiel, Douglas P. http://orcid.org/0000-0001-8474-0310
Kennedy, Brian K. http://orcid.org/0000-0002-5754-1874
Cummings, Steven http://orcid.org/0000-0001-8808-260X
Slagboom, P. Eline http://orcid.org/0000-0002-2875-4723
Verdin, Eric http://orcid.org/0000-0003-3703-3183
Maier, Andrea B.
Sebastiano, Vittorio
Snyder, Michael P. http://orcid.org/0000-0003-0784-7987
Gladyshev, Vadim N. http://orcid.org/0000-0002-0372-7016
Horvath, Steve http://orcid.org/0000-0002-4110-3589
Ferrucci, Luigi http://orcid.org/0000-0002-6273-1613
Article History
Received: 7 September 2023
Accepted: 19 December 2023
First Online: 14 February 2024
Competing interests
: M.M., V.S., M.P.S. and V.N.G. have filed a patent on measuring cellular aging. C.H. is also affiliated with the Institute for Biomedical Aging Research, Universität Innsbruck, Austria and is an honorary research fellow at the Department of Women’s Cancer, EGA Institute for Women’s Health, University College London. C.H. is a shareholder of Sola Diagnostics and is named as an inventor on a patent on an epigenetic clock indicative of breast cancer risk. J.N.J. is also affiliated with the Sticht Center for Healthy Aging and Alzheimer’s Prevention, Wake Forest University School of Medicine and the XPRIZE Foundation. J.N.J. serves on the advisory board for the American Federation for Aging Research’s Finding Aging Biomarkers by Searching Existing Trials Initiative and the editorial board of the <i>Journals of Gerontology Series A Biological Sciences</i>, <i>eLife</i> and <i>Experimental Gerontology</i>. D.W.B. is also affiliated with the Child Brain Development Network, Canadian Institute for Advanced Research and SocioMed Research Nucleus and Universidad Mayor. D.W.B. is an inventor of DunedinPACE, a Duke University and University of Otago invention licensed to TruDiagnostic. A.T.H.-C. is an inventor of epigenetic clocks that are the subject of a provisional patent and have been licensed to TruDiagnostic. A.T.H.-C. has also received consulting fees from TruDiagnostic and FOXO Biosciences. B.H.C. owns stock in Illumina, the manufacturer of the DNA methylation arrays used in epigenetic biomarkers of aging, and is listed as a co-inventor on filed patents on commercial applications of epigenetic prediction models. A.A.C. is a founder, president and majority shareholder at Oken Health. R.E.M. has received a speaker fee from Illumina and is an advisor to the Epigenetic Clock Development Foundation and Optima Partners. M.W. is also affiliated with the Institute for Biomedical Aging Research, Universität Innsbruck. M.W. is a shareholder of Sola Diagnostics and is named as an inventor on a patent on an epigenetic clock indicative of breast cancer risk. K.F. is the CEO of BioAge Labs. P.O.F. is an employee and stakeholder of Gero. A.Z. is the founder and the CEO of Insilico Medicine, a clinical-stage generative AI and robotics biotechnology company specializing in aging research. N.B. is the scientific director of the American Federation for Aging Research, is on the board of the executive committee of the Longevity Biotech Association and is advisor on the board of the Academy for Health and Lifespan Research. D.P.K. has received a grant from Solarea Bio and royalties from Wolters Kluwer. D.P.K. sits on the scientific advisory boards of Solarea Bio, Pfizer, Radius Health and Reneo and has participated in the data safety monitoring board for the AgNovos Healthcare treatment trial. E.V. is a scientific cofounder of Napa Therapeutics and BHB Therapeutics, serves on the scientific advisory board of Seneque and is a named co-inventor on a patent relating to an epigenetic clock robust to cell composition changes. A.B.M. declares herself chief medical officer of NU and co-founder of Chi Longevity. V.S. is a cofounder, SAB chair and head of research of Turn Biotechnologies. M.P.S. is a cofounder and scientific advisor of Personalis, SensOmics, Qbio, January AI, Fodsel, Filtricine, Protos, RTHM, iollo, Marble Therapeutics, Crosshair Therapeutics and Mirvie. He is a scientific advisor of Jupiter, Neuvivo, Swaza and Mitrix. S. Horvath is a founder of the nonprofit Epigenetic Clock Development Foundation that licenses patents surrounding epigenetic clocks. The Regents of the University of California is the sole owner of a patent application directed at GrimAge and other epigenetic clocks for which S. Horvath is a named inventor.