Goosen, Wynand J.
Kerr, Tanya J.
Kleynhans, LĂ©anie
Warren, Robin M.
van Helden, Paul D.
Persing, David H.
Parsons, Sven D. C.
Buss, Peter
Miller, Michele A.
Funding for this research was provided by:
Harry Crossley Foundation
South African Medical Research Council (86949)
American Association of Zoo Veterinarians (S005651)
National Geographic Society (NGS-61089C-19)
Article History
Received: 3 October 2019
Accepted: 17 August 2020
First Online: 2 September 2020
Competing interests
: The authors declare no competing interests.