Li, Juan
Zhang, Na
Zhang, Ziyin
Fu, Jinsai
Ren, Wenjing
Sun, Yi
Song, Shuling
Liu, Xiaoqian
Liu, Jinghui
Wang, Jingyu
Sun, Yunliang
Zhang, Kai
Guo, Rongrong
Lv, Changjun
Pan, Lei
Qu, Guiwu
Han, Fang
Yu, Yan
Funding for this research was provided by:
Natural Science Foundation of Shandong Province (ZR2020MH131)
Natural Science Foundation of Shandong Province (ZR2021MH360)
National Natural Science Foundation of China (82200109)
National Natural Science Foundation of China (82370092)
Youth Innovation and Technology Support Program in Colleges and Universities of Shandong Province (2022KJ090)
Scientific Research Foundation of Fang Han Team
Article History
Received: 26 February 2025
Accepted: 29 August 2025
First Online: 29 October 2025
Change Date: 30 May 2026
Change Type: Update
Change Details: The original online version of this article was revised: The author Juan Li, Na Zhang, Ziyin Zhang, and Jinsai Fu should have been denoted as an equally contributing authors and co-first authors. It has been updated.
Change Date: 8 June 2026
Change Type: Correction
Change Details: A Correction to this paper has been published:
Change Details: https://doi.org/10.1186/s10020-026-01519-z
Declarations
:
: The study was conducted following the guidelines set by the National Institutes of Health’s Guide for the Care and Use of Laboratory Animals. The animal experiments were approved by the Ethics Committees on Animal Experimentation of Binzhou Medical University (No. 2022 − 202).
: Not applicable.
: The authors declare no competing interests.