Alter, S. Elizabeth https://orcid.org/0000-0003-0597-9206
Chiozzi, Giorgio https://orcid.org/0000-0002-1811-0254
De Marchi, Giuseppe https://orcid.org/0000-0002-5894-9089
Fasola, Mauro https://orcid.org/0000-0002-7366-5893
Kurata, Naoko P. https://orcid.org/0000-0002-4236-2646
Stiassny, Melanie L. J. https://orcid.org/0000-0001-8220-4768
Article History
Received: 21 October 2025
Accepted: 11 March 2026
First Online: 2 April 2026
Declarations
:
: This study adhered to ethical guidelines for research involving animals and was approved by the University of Pavia as part of the research programme of the Doctoral School in Earth and Environmental Sciences of one of the authors (GC). The Italian Ministry of the Environment, Land and Sea, as the national authority responsible for implementing the Convention on Biological Diversity, supported the project in the context of international relations (Ref. No. 0025849/PNM, 24 December 2015). Collection of fish specimens was carried out under the appropriate permits issued by the Ministry of Marine Resources and the Ministry of Agriculture of Eritrea, and the Ethiopian Biodiversity Institute (Ref. No. EBI71/2354/2016) in full compliance with institutional, national, and international ethical and legal standards.
: Not applicable.
: Not applicable.
: The authors declare no competing interests.