Mao, Jingyi
He, Junmin
Ren, Yifan
Li, Xue
Yang, Cunming
Liu, Guifen
Zhang, Guoping
Wei, Chen
Zhang, Wenhao
Wang, Ming
Nie, Fumei
Tian, Kechuan
Huang, Xixia
Funding for this research was provided by:
the Agricultural Scientific and Technological Innovation Project of Shandong Academy of Agricultural Sciences (No. CXGC2025F09, No. CXGC2025H16)
the Agricultural Scientific and Technological Innovation Project of Shandong Academy of Agricultural Sciences (No. CXGC2025F09, No. CXGC2025H16)
the Agricultural Scientific and Technological Innovation Project of Shandong Academy of Agricultural Sciences (No. CXGC2025F09, No. CXGC2025H16)
the Agricultural Scientific and Technological Innovation Project of Shandong Academy of Agricultural Sciences (No. CXGC2025F09, No. CXGC2025H16)
the Agricultural Scientific and Technological Innovation Project of Shandong Academy of Agricultural Sciences (No. CXGC2025F09, No. CXGC2025H16)
the Agricultural Scientific and Technological Innovation Project of Shandong Academy of Agricultural Sciences (No. CXGC2025F09, No. CXGC2025H16)
the Agricultural Scientific and Technological Innovation Project of Shandong Academy of Agricultural Sciences (No. CXGC2025F09, No. CXGC2025H16)
the Agricultural Scientific and Technological Innovation Project of Shandong Academy of Agricultural Sciences (No. CXGC2025F09, No. CXGC2025H16)
the Agricultural Scientific and Technological Innovation Project of Shandong Academy of Agricultural Sciences (No. CXGC2025F09, No. CXGC2025H16)
the Agricultural Scientific and Technological Innovation Project of Shandong Academy of Agricultural Sciences (No. CXGC2025F09, No. CXGC2025H16)
the Agricultural Scientific and Technological Innovation Project of Shandong Academy of Agricultural Sciences (No. CXGC2025F09, No. CXGC2025H16)
the Agricultural Scientific and Technological Innovation Project of Shandong Academy of Agricultural Sciences (No. CXGC2025F09, No. CXGC2025H16)
the Technological Innovation Project of the National Natural Science Foundation of China (No. 32372851)
the Technological Innovation Project of the National Natural Science Foundation of China (No. 32372851)
the Technological Innovation Project of the National Natural Science Foundation of China (No. 32372851)
the Technological Innovation Project of the National Natural Science Foundation of China (No. 32372851)
the Technological Innovation Project of the National Natural Science Foundation of China (No. 32372851)
the Technological Innovation Project of the National Natural Science Foundation of China (No. 32372851)
the Technological Innovation Project of the National Natural Science Foundation of China (No. 32372851)
the Technological Innovation Project of the National Natural Science Foundation of China (No. 32372851)
the Technological Innovation Project of the National Natural Science Foundation of China (No. 32372851)
the Technological Innovation Project of the National Natural Science Foundation of China (No. 32372851)
the Technological Innovation Project of the National Natural Science Foundation of China (No. 32372851)
the Shandong Postdoctoral Science Foundation (No. SDCX-ZG-202400120)
the Shandong Postdoctoral Science Foundation (No. SDCX-ZG-202400120)
the Shandong Postdoctoral Science Foundation (No. SDCX-ZG-202400120)
the Shandong Postdoctoral Science Foundation (No. SDCX-ZG-202400120)
the Shandong Postdoctoral Science Foundation (No. SDCX-ZG-202400120)
the Shandong Postdoctoral Science Foundation (No. SDCX-ZG-202400120)
the Shandong Postdoctoral Science Foundation (No. SDCX-ZG-202400120)
Article History
Received: 3 June 2025
Accepted: 29 October 2025
First Online: 15 December 2025
Declarations
:
: All animal experiments were approved by the Laboratory Animal Ethics Committee, Institute of Animal Science and Veterinary Medicine, Shandong Academy of Agricultural Sciences (Jinan, China, IASVM-2022-003).
: Not applicable.
: The authors declare no competing interests.